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Arrowhead Research
ARWR

Name Holding Trade Value Shares
Change 		Change in
Stake
ARTA
226
A.R.T. Advisors
New York
– -$1.52M -57,325 Closed
Voloridge Investment Management
227
Voloridge Investment Management
Florida
– -$1.43M -53,790 Closed
DekaBank Deutsche Girozentrale
228
DekaBank Deutsche Girozentrale
Germany
– -$1.11M -38,400 Closed
Jacobs Levy Equity Management
229
Jacobs Levy Equity Management
New Jersey
– -$888K -33,500 Closed
PCM
230
Penserra Capital Management
California
– -$790K -29,843 Closed
VRS
231
Virginia Retirement Systems
Virginia
– -$610K -23,000 Closed
USAA
232
United Services Automobile Association
Texas
– -$544K -20,515 Closed
CCIA
233
Campbell & Co Investment Adviser
Maryland
– -$389K -14,679 Closed
WG
234
Winton Group
United Kingdom
– -$367K -13,836 Closed
KIG
235
Kessler Investment Group
Indiana
– -$355K -13,380 Closed
WCM
236
Weld Capital Management
New York
– -$289K -10,910 Closed
HF
237
HRT Financial
New York
– -$269K -10,173 Closed
Mariner
238
Mariner
Kansas
– -$264K -9,971 Closed
JMPWA
239
JP Morgan Private Wealth Advisors
California
– -$227K -8,564 Closed
CIP
240
Convergence Investment Partners
Florida
– -$186K -7,015 Closed
QT
241
Quantamental Technologies
New York
– -$156K -5,886 Closed
PRC
242
Pearl River Capital
Michigan
– -$111K -4,188 Closed
MWM
243
Mascoma Wealth Management
New Hampshire
– -$46K -1,745 Closed
CAW
244
Cornerstone Advisors (Washington)
Washington
– -$38K -1,446 Closed
AC
245
Avestar Capital
New York
– -$10K -413 Closed
OIA
246
OneDigital Investment Advisors
Kansas
– -$11K -400 Closed
FPFS
247
First Personal Financial Services
North Carolina
– -$6K -218 Closed
TCM
248
Tradewinds Capital Management
Washington
– -$5K -200 Closed
PCP
249
Proficio Capital Partners
Massachusetts
– -$5K -188 Closed
CFGS
250
Concourse Financial Group Securities
Alabama
– -$4K -135 Closed
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